Research funding: This research was supported by the Key Laboratory of Bio-based Material Science & Technology (Northeast Forestry University), Ministry of Education of the People’s Republic of China (SWZ-MS201917), Hunan Provincial Technical Innovation Platform and Talent Program in Science and Technology, PR China (2019RS2040), Major Science and Technology Program of Hunan Province, PR China (2017NK1010), National Natural Science Foundation of China (31770606), Hunan Province Innovation Foundation for Postgraduate, PR China (CX20190600) and Science and Technology Innovation Project for Postgraduates of Central South Forestry University of Science and Technology, PR China (CX20191010, CX20192003).