All authors contributed to this study’s conception and design. Investigation, writing—original draft preparation, and writing—review and editing, W.L.; supervision and validation, H.Y.; data curation and formal analysis, H.W.; conceptualization and funding acquisition, X.J.; resources, Z.L. All authors commented on previous versions of the manuscript. All authors have read and agreed to the published version of the manuscript. This study was supported by the Special Projects in Key Fields of Higher Education in Guangdong Province (Grant No.: 2023ZDZX4044), the Special Projects in Key Fields of Higher Education in Guangdong Province (Grant No.: 2023ZDZX4045), the Research Capacity Enhancement Project of Key Construction Discipline in Guangdong Province (Grant No.: 2022ZDJS092), the National Natural Science Foundation of China (31971727), and the Forest Science and Technology Innovation Program of Hunan Province (XLK202105-3).