Conceptualization, Methodology, Software, Investigation, Methodology, Writing—Original draft preparation, Writing—Reviewing and Editing, G.L.; Conceptualization, Methodology, Software, Investigation, Methodology, C.L.; Conceptualization, Methodology, Software, Investigation, Methodology, G.J.; Conceptualization, Methodology, Software, Investigation, Methodology, Z.H.; Writing—Original draft preparation, Writing—Reviewing and Editing, Funding, Y.H.; Investigation, Data curation, Formal analysis, Supervision, Project administration, Funding acquisition, Writing—Reviewing and Editing, W.H.; The authors G.L. and C.L. have made equal contributions. All authors have read and agreed to the published version of the manuscript. This research was funded by the Natural Science Foundation of Hunan Province [Grant numbers:2022JJ40862], the Key Project of Hunan Education Department [Grant number: 21A0513], the Scientific Research Project of Hunan Education Department [Grant numbers 21B0235] and the Natural Science Foundation of Hunan Province [Grant number: 2020JJ4942]. This work was supported by the Key Discipline of the State Forestry Administration [Grant number: 2016-21] and the “Double First-Class” Cultivating Subject of Hunan Province [Grant number: 2018-469].