Conceptualization, L.L. and P.Y.; methodology, L.L.; software, H.L.; validation, P.Y. and H.L.; investigation, P.Y. and Y.H.; resources, L.L.; writing—original draft preparation, L.L.; writing—review and editing, M.Z. and K.B.B.; supervision, K.B.B. and H.L.; funding acquisition, L.L. All authors have read and agreed to the published version of the manuscript. This research was funded by the National Natural Science Foundation of China, grant number 31901363, the Scientific Innovation Fund for Postgraduates of Central South University of Forestry and Technology, grant number CX202202076, and the Outstanding Youth Project of Hunan Provincial Department of Education, grant number 22B0285.