[He, Gong-Xiu;Zhang, Dang-Quan] Cent S Univ Forestry & Technol, Hunan Prov Key Lab Forestry Biotechnol, Changsha 410004, Hunan, Peoples R China.^[Zhang, Dang-Quan;Guo, Lin-Lin] Cent S Univ Forestry & Technol, Key Lab Non Wood Forest Product State Foresty Adm, Biotechnol Core Facil, Changsha 410004, Hunan, Peoples R China.