Conceptualization, S.L.; methodology, S.L. and J.Z.; validation, Z.W., S.F. and L.J.; investigation, J.Z.; writing—original draft preparation, J.Z.; writing—review and editing, S.L., H.G., X.L., B.W., Z.H. and. F.X.; visualization, Z.W., S.F. and Z.H.; funding support, S.L. and X.L. All authors have read and agreed to the published version of the manuscript. This research was funded by the National Natural Science Foundation of China (Grant No. 41971152 and U20A2089) to S.L.; and the Hunan Provincial Forestry Science and Technology Innovation Fund Project (Grant No. XLK202103-2) to X.L.