The authors gratefully acknowledge financial support by the Technology Foundation for Selected Overseas Chinese Scholar of Ministry of Hunan Resources and Social Security of China (2016-176), the Hunan Provincial Key Laboratory Special Foundation of China (2014FJ3011, 2016XGJSYB01 and 2015TP4021-5), the Natural Science Foundation of Hunan Province, China (2018JJ2671), and Scientific Innovation Fund for Post-graduates of Central South University of Forestry and Technology (20183009).