CRediT authorship contribution statement Junjie Lei: Writing – review & editing, Writing – original draft, Visualization, Software, Investigation, Data curation. Xiaoqian Gao: Writing – original draft, Validation, Supervision, Investigation, Data curation. Ting He: Validation, Investigation, Data curation. Zongxin Liu: Validation, Investigation, Data curation. Wende Yan: Resources, acquisition. Peng Dang: Writing – review & editing, Project administration, acquisition, Formal analysis, Conceptualization. This research was funded by the National Natural Science Foundation of China ( 32001303 ), the Hunan Province key research and development project ( 2023SK2055 ), the China Postdoctoral Science Foundation ( 2020M672524 ), the Hunan Provincial Natural Science Foundation of China ( 2024JJ5639 ), and the Changsha Municipal Natural Science Foundation ( kq2402249 ).