Conceptualization, Y.Z., H.G., H.D. and Y.H.; methodology, Y.Z., H.G. and H.D.; software, Y.Z.; validation, H.G. and H.D.; formal analysis, Y.Z. and H.G.; investigation, J.Y. and J.W.; resources, H.G.; data curation, Y.Z. and H.G.; writing—original draft preparation, Y.Z.; writing—review and editing, H.G., H.D., Z.X. and Y.H.; visualization, Y.Z.; supervision, Z.X. and Y.H.; project administration, Z.X. and Y.H.; funding acquisition, Z.X. and Y.H. All authors have read and agreed to the published version of the manuscript. This study was supported by the Forestry Science and Technology Innovation Project of Guangdong Province (2023KJCX023), Science and Technology Project of Guangzhou, Grant/Award Number: 2023A04J0836.