Conceptualization, S.L. and S.Z.; methodology, S.L.; software, H.X.; validation, S.Z., H.X. and Z.G.; formal analysis, S.Y.; investigation, Z.G.; resources, S.Z.; data curation, S.Y.; writing—original draft preparation, S.L.; writing—review and editing, S.Z.; visualization, H.X.; supervision, S.Z.; project administration, S.Z.; funding acquisition, S.Z. All authors have read and agreed to the published version of the manuscript. This study was financially supported by the Forestry Public Welfare Scientific Research Project of China, funding number 201504301.