Conceptualization, Z.L. and X.H.; methodology, Z.L.; software, Z.L.; validation, Z.L. and Y.W.; formal analysis, Z.L.; investigation, Z.L.; resources, Z.L.; data curation, Z.L.; writing—original draft preparation, Z.L.; writing—review and editing, Z.L., X.H., Y.W. and C.C.; visualization, Z.L.; supervision, X.H.; project administration, X.H.; funding acquisition, X.H. All authors have read and agreed to the published version of the manuscript. This research was funded by the Key Disciplines of State Forestry Administration of China, grant number No. 21 of Forest Ren Fa, 2016; Hunan Province “Double First-class” Cultivation discipline of China, grant number No. 469 of Xiang Jiao Tong, 2018; and Hunan Provincial Innovation Foundation for Postgraduate, grant number CX20210850.