This work was supported by the National Key Research and Development Program of China (Grant number 2017YFD0601104), the Project for Cultivating Excellent Doctoral Dissertation of Forestry Engineering (Grant number LYGCYB202001), the Scientific Innovation Fund for Post-graduates of Hunan Province (Grant number QL20210208), the Scientific Innovation Fund for Post-graduates of Central South University of Forestry and Technology (Grant number CX202101022).