Conceptualization, P.H., W.Y. and X.C.; methodology, P.H., W.Y., Y.P. and X.C.; validation, J.L., W.Z., Y.Z. and Y.Q.; formal analysis, Y.P., J.L., W.Z., Y.Z. and Y.Q.; investigation, P.H., J.L. and W.Z.; resources, J.L., Y.Z. and Y.Q.; writing—original draft preparation, P.H. and Y.P; writing—review and editing, Y.P., J.L. and X.C.; supervision, W.Y. and X.C.; project administration, P.H. and W.Y.; funding acquisition, W.Y. and X.C. All authors have read and agreed to the published version of the manuscript. The study was supported by the National Key Research and Development Program of China (2020YFA0608101), the Joint Funds of the National Natural Science Foundation of China (U21A20187), and a ‘Shu Ren Scholar’ plan of Central South University of Forestry and Technology. We would like to thank Cao Zhang, Wancai Wang, and Dongjun Zou for their field and laboratory measurements.