Acknowledgments As a key project under the State Forestry Administration of China (2006-11, 2006-17, 2005-08), this project was funded by the National Natural Science Foundation of China (No. 30571487, 30771700), the Furong Scholar Program, the Urban Forest Ecological Key Laboratory of Hunan Province (No. 06FJ3083), and the Platform Construction Project under the Ministry of Science and Technology of China.