This work was supported by the: Special Fund for Forest Scientific Research in the Public Welfare , under Grant number 201404314 ; China Scholarship Council Foundation , under Grant number 201408430015 ; China Postdoctoral Science Foundation , under Grant number 2014M550887 ; China National Tourism Administration Young Expert Fund , under Grant [number TYEPT201437 ]; Hunan Province Philosophy and Social Science Fund , under Grant number 14JD68 ; and Hunan Province Education Science Twelfth-Five Planning Fund , under Grant number XJK011QXJ003 .