CRediT authorship contribution statement Hui Chen: Writing – original draft, Visualization, Methodology, Investigation, Formal analysis, Conceptualization. Ying Zhang: Investigation, Formal analysis, Data curation. Yi Tan: Formal analysis, Data curation. Jiawei Shao: Formal analysis. Xinyi Li: Formal analysis. Mingyang Bai: Formal analysis. Shanshan Gong: Writing – review & editing. Tao Liu: Writing – review & editing, Supervision, Formal analysis. Jianzhang Li: Writing – review & editing, Supervision, acquisition. Acknowledgments The authors acknowledge the funding supported by the 5·5 Engineering Research & Innovation Team Project of Beijing Forestry University ( BLRC2023A02 ), National Natural Science Foundation of China ( 32101600 ) and National Natural Science Foundation of China ( 32201641 ).