This work was financially supported by the Natural Science Foundation of Hunan Province (No. 2023JJ31011), the Natural Science Foundation of Changsha City (No. kq2202276), the National Natural Science Foundation of China (No. 51608208), the Natural Science Foundation of Hunan Province (No. 2019JJ51005), the Key Research and Development Program of Hunan Province (No. 2019SK2191), the Natural Science Foundation of Hunan Province (No. 2020JJ5984), and the Scientific Innovation Fund for Graduate of Central South University of Forestry and Technology (CX20192010).