National Natural Science Foundation of ChinaNational Natural Science Foundation of China [21908251, 31890771, 31870552]; Hunan Provincial Natural Science Foundation of ChinaNatural Science Foundation of Hunan Province [2017JJ1038, 2019JJ40535]; National Key Research and Development Program of China [2017YFD0600202]; PhD research startup foundation of Central South University of Forestry and Technology [104-0456]; Youth Scientific Research Foundation, Central South University of Forestry and Technology [QJ2018003B]; Hunan high-level talent gathering project-innovative talents [2019RS1061]