This study was supported by the Key Research and Development Program of Hunan Province (2021NK2027), the National Natural Science Foundation of Hunan Province (2021JJ31159 and 2022JJ40872), the Scientific Research Project of Education Department of Hunan Province of China (21B0223), Changsha Municipal Natural Science Foundation (kq2202287), and the Introduced Talent Research Startup Fund of Central South University of Forestry and Technology (2021YJ020).