All authors read the manuscript; conceptualization and methodology, H.S. and F.Z.; validation, F.J. and F.Z.; formal analysis, F.J.; investigation, F.J., D.L., and H.S.; draft, F.J., H.S., and K.M.; supervision, H.S.; review, editing, and revision, F.Z. and K.M.; funding acquisition, H.S. All authors have read and agreed to the published version of the manuscript. This research was funded by the project of the National Natural Science Foundation of China, grant number 31971578; Scientific Research Fund of Hunan Provincial Education Department, grant number 17A225; National Bureau to Combat Desertification, State Forestry Administration of China, grant number 101-9899; and the Forestry Administration of Hunan Province, grant number XLK201986.