Fundings This work was supported by the Forestry Science and Technology Innovation and Promotion Project of Jiangsu Province ( LYKJ[2024]08 ), the National Natural Science Foundation of China (No. 51878354 & 51308301 ), the Natural Science Foundation of Jiangsu Province (No. BK20181402 & BK20130978 ). All research outcomes presented in this paper are those of the writer(s) and do not necessarily reflect the views of the foundations. CRediT authorship contribution statement Linji Ying: Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Resources, Validation, Visualization, Writing - original draft. Haitao Li: Conceptualization, Formal analysis, acquisition, Investigation, Supervision, Writing - original draft. Yiqiang Wu: Investigation, Validation, Writing - review. Tianyu Gao: Investigation, Validation, Writing - review. Wenjing Zhou: Investigation, Writing - review. Ottavia Corbi: Investigation, Writing - review.