W.C. and F.Z. conceived the experiment and developed the experimental design. W.C.; L.H., and S.T. developed and carried out the specific methodology. W.C. and L.H. contributed to the data analysis. D.Y. secured funding for the research and was responsible for overall project administration. F.Z., D.Y. and R.Z. supervised the research of W.C. W.C., L.H.; F.Z., and J.M. contributed to the preparation of the manuscript. All authors have read and agreed to the published version of the manuscript. This research was funded bythe National Natural Science Foundation of China (grant no. 31870674, no.31500554) and the ‘12th Five year’ Science and Technology Support Program of China (grant no. 2013BAD14B04).