The authors gratefully acknowledge the financial support of the National Natural Science Foundation of China (Grant Nos: 41977129 , 21607176 ), Hunan Huxiang Young Talents Support Program , China (No. 2020RC3044 ), Changsha Outstanding Innovative Youth Training Program (Grant No. kq1802011 ), the Natural Science Foundation of Hunan Province , China (Grant No. 2020JJ4136 , 2022JJ90020 ), the Research Foundation of Education Bureau of Hunan Province , China (Grant No. 16B274 ), and the Innovation Fund for Post-graduates of Central South University of Forestry and Technology (Grant No. CX20192023 ). The authors also thank the reviewers for their constructive comments and suggestions.