This research was financially supported by the Key Fields of Higher Education in Guangdong Province (Grant Nos. 2023ZDZX4044, 2023ZDZX4045); Research Capacity Enhancement Project of Key Construction Discipline in Guangdong Province (Grant No. 2022ZDJS092); project supported by National Natural Science Foundation of China (Grant No. 31971727); The Forest Science and Technology Innovation Program of Hunan Province (Grant No. XLK202105-3); Hunan Provincial Natural Science Foundation of China (Grant No. 2022JJ31005).