This work was financially supported by the Major Science and Technology Program of Hunan Province (2017NK1010), National Natural Science Foundation of China (31870552), Key Laboratory of Bio-based Material Science & Technology, Ministry of Education (SWZCL2016-02), Young Elite Scientists Sponsorship Program by CAST (2016QNRC001), Hunan Provincial Technical Innovation Platform and Talent Program in Science and Technology (2016RS2010, 2016TP1013), and Science and Technology Innovation Fund for Graduate Students, from Central South University of Forestry and Technology (20181013).