Methodology, Q.L. and Y.L.; formal analysis, Q.L., Y.H., Y.L., C.C. and Y.H.; investigation, Z.Y. and Y.Z.; data curation, Q.L. and J.L.; writing—original draft preparation, Q.L. and C.C.; project administration, Y.L. and J.L.; funding acquisition, Q.L. and J.L. All authors have read and agreed to the published version of the manuscript. This research was funded by the Hunan Forestry Science and Technology Innovation Project of China (Grant No. XLKT202303), the Forest Science and Technology Innovation Project of Hunan Province (Grant No. XLKJ202203), the Hunan Forestry Science and Technology Innovation Project (Grant No. XLKY202313), and the Key Scientific Research Project of Hunan Provincial Department of Education (Grant No. 22A0197).