Y.C.: conceptualization, methodology, investigation, formal analysis, and writing—review and editing. X.G.: investigation, writing—original draft, and validation. H.X.: formal analysis. Q.W., A.X., Z.Z. and R.T.: investigation. W.G.: supervision, writing—review and editing, and funding acquisition. D.Y.: supervision, resources, and funding acquisition. All authors have read and agreed to the published version of the manuscript. This research was funded by the Science and Technology Innovation Program of Hunan Province (2022RC1152), the National Natural Science Foundation of China (grant 32371931), the Forestry Science and Technology Innovation Program of Hunan Province (XLK202101-2), the major science and technology project of Changsha Bureau of Science and Technology (kq2102007), and the Scientific Research Foundation for Advanced Talents of Central South University of Forestry and Technology (2018YJ002).
