Conceptualization, Data curation, Formal analysis, Methodology, Visualization, Writing—original draft, Writing—review and editing: Q.W.; Formal analysis: Y.L.; Visualization: T.X. and H.L.; Formal analysis, Writing—review and editing: Z.H. and Z.C.; Conceptualization, Data curation, Formal analysis, Methodology, Writing—original draft, Writing—review and editing, Funding acquisition, Supervision: F.L. All authors have read and agreed to the published version of the manuscript. The authors gratefully acknowledge the funds from the National Key R&D Program of China (No. 2022YFF1100200), the Natural Science Foundation of Hunan Province, China (Grant No. 2021JJ31075), the Program for Science & Technology Innovation Platform of Hunan Province (2019TP1029), Hunan Provincial Innovation Foundation For Postgraduate (No. CX20230765), Scientific Innovation Fund for Post-graduates of Central South University of Forestry and Technology (No. 2023CX02025).