Conceptualization, N.Z. and P.D.; Methodology, P.D.; Validation, X.G. and Z.L.; Formal analysis, L.L.; Data curation, L.L. and T.H.; Writing—original draft, L.L. and T.H.; Writing—review & editing, Y.P. and P.D.; Funding acquisition, P.D. All authors have read and agreed to the published version of the manuscript. This research was funded by the National Natural Science Foundation of China (32001303), the Changsha Municipal Natural Science Foundation (kq2014155), Forestry science and technology research and innovation project of Hunan Forestry Bureau (XLKY202330); the China Postdoctoral Science Foundation (2020M672524).