This work was financially supported by National Natural Science Foundation of China ( 31890771 and 32201646 ), Young Elite Scientists Sponsorship Program from National Forestry and Grassland Administration of China ( 2019132614 ), Hunan Provincial Technical Innovation Platform and Talent Program in Science and Technology ( 2018RS3092 and 2018WK4028 ), and Outstanding Innovative Youth Training Program of Changsha ( KQ2009059 ).