Conceptualization, L.J., L.W., Q.Z. and C.L.; data curation, L.W.; formal analysis, G.L. and L.W.; funding acquisition, W.L. and G.L.; investigation, L.J., K.H., P.L. and R.F.; methodology, L.J. and L.W.; project administration, W.L. and C.L.; software, L.W.; supervision, L.W.; validation, L.J., G.L. and C.L.; visualization, L.J. and C.L.; writing—original draft, L.J., Q.Z. and C.L.; writing—review and editing, L.W. and C.L. All authors have read and agreed to the published version of the manuscript. This research was funded by the National Key Research and Development Program of China, grant number 2017YFC0405300; the National Natural Science Foundation of China, grant numbers U20A2089 and 41971152; and the Natural Science Foundation of Hunan Province, China, grant number 2021JJ20012.