This study was supported by the Hunan Provincial Technical Innovation Platform and Talent Program in Science and Technology (grant no. 2020RC3041), the Training Program for Excellent Young Innovators of Changsha (grant no. kq2009015), the Hunan Provincial Natural Science Foundation of China (grant no. 2022JJ30079 ), the China Postdoctoral Science Foundation (Grant Nos. 2020M672846 and 2022T150556 ), and the Postgraduate Technology Innovation Project of Central South University of Forestry and Technology (2023CX02073).