Conceptualization, L.Z. and F.L.; methodology, L.Z., Z.H., Y.Z. and T.G.; validation, L.Z., Z.H., Y.Z. and T.G.; formal analysis, L.Z.; investigation, L.Z., Z.H., Y.Z., T.G., Q.X. and H.Y.; resources, F.C., Q.L. and F.L.; data curation, L.Z., Z.H., Y.Z., T.G., S.Y. and X.X.; writing—original draft preparation, L.Z.; writing—review and editing, F.L.; visualization, L.Z.; supervision, F.L; project administration, L.Z. and F.L.; funding acquisition, G.L. All authors have read and agreed to the published version of the manuscript. This study was financially supported by Hunan Province Modern Agricultural Industrial Technology System-Dry Grain Industrial Technology System project, Agricultural Science Technology Innovation Fund project of Hunan Province (2022CX27) and Changsha Natural Science Foundation project (kq2202333), Hunan Province, China.