Q.G., F.L. and Y.D.: conceptualization, supervision, funding acquisition, writing—review and editing. M.H.: investigation, sample collection, data acquisition, formal analysis, writing—original manuscript. Y.Y. and S.G.: sample collection, methodology, validation, formal analysis. H.S., Q.L. and X.L.: validation, writing—review and editing. All authors have read and agreed to the published version of the manuscript. This work was supported by the Science and Technology Innovation Program of Hunan Province (2022RC1032, 2023JJ40644), Key R & D Program of Hunan Province (2022NK2026), and China Agriculture Research System of MOF and MARA (CARS-35).