This work was supported by National Science Foundation of China (No. 31890772 ), the Natural Science Foundation of Hunan Province ( 2021JJ41076 ), the Science and Technology Innovation Program of Hunan Province ( 2020RC4049 ), the Excellent Youth Project of Scientific Research Project of Hunan Provincial Department of Education ( 19B603 ), the Central South University of Forestry Science and Technology Introduced Talents Scientific Research Startup Fund Project ( 2019TJ014 ). Professor Goodell was also supported by the National Institute of Food and Agriculture , U.S. Department of Agriculture , the Center for Agriculture, Food and the Environment , and the Microbiology department at University of Massachusetts Amherst , under project number S1075 - MAS00503 . The contents are solely the responsibility of the authors and do not necessarily represent the official views of the USDA or NIFA.