Conceptualization, Z.Z.; methodology, J.Z.; validation, Z.G., data curation, Y.C. and L.X.; writing—original draft preparation, J.Z.; writing—review and editing, F.S. and Z.G.; visualization, Y.C. and Z.G.; funding acquisition, J.Z. All authors have read and agreed to the published version of the manuscript. This research was funded by Chinese national Promotion Program of Forestry and Grassland Scientific and Technological Achievements, grant number 2020133139.