Conceptualization, Z.L., Q.W., S.L., W.Y. and X.C.; methodology, Z.L., Y.P. and W.Y.; validation, Z.L., Q.W., C.M. and X.L.; formal analysis, J.L., Y.P. and J.W.; investigation, Z.L., Q.W., J.L., C.M. and X.L.; resources, J.L., Q.W., C.M. and X.L.; writing—original draft preparation, Z.L., Q.W. and Y.P.; writing—review and editing, Z.L., Y.P., W.Y. and X.C.; visualization, J.L. and J.W.; supervision, S.L., W.Y. and X.C.; project administration, Z.L. and W.Y.; funding acquisition, S.L. and W.Y. All authors have read and agreed to the published version of the manuscript. This study was funded by the National Key R&D Program of China (grant number: 2020YFA0608100), Joint Funds of the National Natural Science Foundation of China (grant number: U21A20187), and the Key Research and Development Program of Hunan Province (2020NK2022).