All authors have read and agreed to the published version of the manuscript. Conceptualization, H.S. and G.W.; methodology, H.S. and G.W.; validation, C.L.; formal analysis, C.L.; investigation, Y.C., H.S., and X.X.; writing—original draft, Y.C., H.S., and C.L.; supervision, H.S.; writing—review and editing, G.W.; project administration, G.W.; funding acquisition, H.S. All authors have read and agreed to the published version of the manuscript. This research was funded by the National Bureau to Combat Desertification, State Forestry Administration of China, grant number101-9899; Scientific Research Fund of Hunan Provincial Education Department, grant number 17A225; the project of the National Key R&D Program of China, grant number 2017YFC0506502 and Training Fund of Young Professors from Hunan Provincial Education Department, grant number 90102-7070220090001.