The authors contributed as follow: Conceptualization, Q.T. and D.M.; methodology, Y.Z.; software, Y.Z.; validation, L.W., W.C., and J.W.; formal analysis, Y.Z.; investigation, Y.Z.; resources, X.W.; data curation, D.Q.; writing—original draft preparation, Y.Z. and D.M.; writing—review and editing, Q.T., Z.L. and I.W.W.; visualization, Y.Z.; supervision, Q.T.; project administration, Q.T.; funding acquisition, Q.T. All authors have read and agreed to the published version of the manuscript. This research was funded by the National Natural Science Foundation of China (No. 32270398), Postgraduate Scientific Research Innovation Project of Hunan Province (No. CX20220689) and Postgraduate Scientific Research Innovation Project of Hunan Agriculture University (No. 2022XC062).