Univ Quebec, Dept Sci Biol, Inst Environm Sci, Montreal, PQ H3C 3P8, Canada.^Chinese Acad Sci, Inst Geog Sci & nat Resources Res, Beijing 100101, Peoples R China.^Zhejiang Forestry Univ, Int Res Ctr Spatial Ecol & Ecosyst Ecol, Hangzhou 31130, Peoples R China.^Nanjing Univ, Int Inst Earth Syst Sci, Nanjing 210093, Jiangshu, Peoples R China.^Cent S Univ Forestry & Technol, Coll Life Sci & Technol, Changsha 410004, Peoples R China.^Chinese Acad Forestry, Inst Forest Resource Informat Tech, Beijing 100091, Peoples R China.^Canada Ctr Remote Sensing, Ottawa, ON K1A OY7, Canada.