The work was financed by Training Plan of Young Backbone Teachers in Colleges and Universities of Hunan Province , China (Grant No. 71 ), Special Projects of Scientific and Technological Innovation in Hunan Forestry , China (Grant No. XLK201982 ), Natural Science Foundation of Hunan Province , China (Grant No. 2019JJ50981 ), Research Foundation of Education Bureau of Hunan Province , China (Grant No. 18A166 ) and Hunan Provincial College Students' Innovation and Entrepreneurship Training Program (Grant No. 2495 ).