Conceptualization, F.W., Z.F., Z.Y., M.L. and Y.W.; methodology, F.W., Z.F., M.L. and Y.W.; software, F.W.; validation, F.W.; formal analysis, F.W.; investigation, F.W., Z.Y., P.Z., M.L. and Y.W.; writing—original draft preparation, F.W. and Z.F.; writing—review and editing, F.W., Z.F., M.L. and Y.W.; supervision, M.L. and Y.W.; project administration, Y.W.; funding acquisition, Y.W. All authors have read and agreed to the published version of the manuscript. This research was funded by the Natural Science Foundation of China (32171842), National Key Research and Development Project of China (2019YFD1100403), Hunan Graduate Research Innovation Project (Key project, Grant No. CX20240068), and Innovation Project of Central South University of Forestry and Technology (90102-63223068).