Conceptualization, P.X.; methodology, H.Z., M.Z. and N.Z.; software, H.Z. and J.C.; validation, P.X., L.M. and J.C.; formal analysis, P.X. and H.Z.; investigation, H.Z. (field sampling), P.X. (field sampling and measurements), M.Z. (multi-omics analysis), S.S. (phenological observation), Q.L. (multi-omics testing), N.Z. and L.M.; data curation, P.X., H.Z. and J.C.; writing—original draft preparation, P.X., H.Z. and S.S.; writing—review and editing, P.X., M.Z. and Q.L.; visualization, P.X. and S.S.; supervision, N.Z. and Q.L.; project administration, M.Z. and S.S.; funding acquisition, M.Z. and S.S. All authors have read and agreed to the published version of the manuscript. This work was supported by the Hunan Forestry Science and Technology Tackling and Innovation Funding Project #1 under grant LKY202327. Central Finance Forestry Science and Technology Promotion Demonstration Project (XLK202213).