Conceptualization, Q.L. and L.J.; methodology, L.Z. and Y.P.; software, Y.A.; validation, Q.L., P.L. and Y.Y.; formal analysis, L.Z.; investigation, A.S.; resources, J.C. and C.L.; data curation, L.Z.; writing—original draft preparation, L.Z. and Y.A. writing—review and editing, Y.C. and Q.L.; visualization, Q.L.; supervision, L.J. and C.L.; project administration, Q.L.; funding acquisition, Y.Y. All authors have read and agreed to the published version of the manuscript. This work was supported by the Scientific Innovation Fund for Post-graduates of Central South University of Forestry and Technology (2023CX01024), the Natural Science Foundation of Hunan Province (2022JJ31001), the Science Foundation of Education Department of Hunan Province (21B0244), the Changsha Municipal Natural Science Foundation (kq2208098 and kq2208099), the Hunan Province Forestry Science and Technology Research and Innovation Project (XLKY202205), and the Changsha Science and Technology Plan Project Assignment (kq2301004).