This work was financially supported by the National Natural Science Foundation of China (31270602), Key Laboratory of Bio-based Material Science & Technology, Ministry of Education (SWZCL2016-02), Young Elite Scientists Sponsorship Program by CAST (2016QNRC001), Key Laboratory of Wood Science and Technology, Zhejiang Province (2014lygcz001), and Hunan Provincial Technical Innovation Platform and Talent Program in Science and Technology (2016RS2010, 2016TP1013).