Conceptualization, Q.L. and Y.Y.; methodology, Y.A., Y.C. and S.Z.; writing—original draft preparation, Y.A.; writing—review and editing, Q.L.; visualization, W.Z. and D.K.; supervision, J.C., P.L. and C.L.; project administration, Q.L.; funding acquisition, L.J. and Y.Y. All authors have read and agreed to the published version of the manuscript. This work was supported by Science and Technology Innovation Program of Hunan Province (2023NK4298), the Innovation Demonstration Project of Chenzhou City (2022sfq53), the Natural Science Foundation of Hunan Province (2021JJ31143), the Key Scientific and Technological Innovation Platform of Hunan Province (2023PT1002) and the State Key Laboratory of Utilization of Woody Oil Resource (GZKF202202).