Conceptualization, Q.L. and D.L.; methodology, Q.L. and D.L.; software, Q.L.; formal analysis, Q.L. and D.L.; investigation, Q.L. and J.G.; resources, Z.G. and D.L.; writing—original draft preparation, Q.L.; writing—review and editing, D.L. and T.W.; visualization, Q.L. and W.Z.; supervision, Z.G. and D.L.; project administration, T.W.; funding acquisition, T.W. and D.L. All authors have read and agreed to the published version of the manuscript. This research was funded by the Key Research and Development Program of Hunan Province, China (2022NK2048), the Natural Science Foundation for Distinguished Young Scholars of Hunan Province (2023JJ10099), Hunan Forestry Science and Technology Project for Distinguished Young Scholars (XLK202108-7), and Key Scientific Research Project of Education Department of Hunan Province, China (22A0187, 20A515).